Browsing by Author "Herman, Lee H."
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ItemCatalog of the Staphylinidae (Insecta, Coleoptera) : 1758 to the end of the second millennium.([New York] : American Museum of Natural History, 2001) Herman, Lee H.; Smetana, Aleš.This catalog (published in seven parts, all released on the same day) is based on only the published literature for the Staphylinidae. Of the 32 subfamilies, the following 28 are included herein: Apateticinae, Dasycerinae, Empelinae, Euaesthetinae, Glypholomatinae, Habrocerinae, Leptotyphlinae, Megalopsidiinae, Micropeplinae, Microsilphinae, Neophoninae, Olisthaerinae, Omaliinae, Osoriinae, Oxyporinae, Oxytelinae, Phloeocharinae, Piestinae, Protactinae†, Proteininae, Protopselaphinae, Pseudopsinae, Solieriinae, Staphylininae, Steninae, Tachyporinae, Trichophyinae, and Trigonurinae. The Aleocharinae, Paederinae, Pselphinae, and Scaphidiinae are excluded from this edition of the catalog. References to the original citation or description are given for available family-group, genus-group, and species-group names of both extant and extinct forms. The type genus is cited for each family-group name, the type species for each genus-group name, and the type locality for each species-group name. Where appropriate, all subgenera, subspecies, or synonyms are listed for each valid name. Annotated subsequent references are presented for all names. Distributional summaries are given for each valid taxon. Full bibliographic citations are in Part VII. A short historical review, coauthored with Aleš Smetana, follows the Introduction (Part I), with the main focus on biographical sketches that include many photographs. The goal of this catalog is to summarize the current state of knowledge of the family and to stimulate worldwide monographic studies. ItemThe ecology, phylogeny, and taxonomy of Stilicolina (Coleoptera, Staphylinidae, Paederinae). American Museum novitates ; no. 2412(New York, N.Y. : American Museum of Natural History, 1970) Herman, Lee H. ItemEppelsheimius : revision, distribution, sister group relationship (Staphylinidae, Oxytelinae). American Museum novitates ; no. 2764(New York, N.Y. : American Museum of Natural History, 1983) Herman, Lee H."Eppelsheimius is a small genus of beetles that occurs in arid regions from northern Africa to southwestern Asia. The species share characters with Planeustomus, Manda, and Bledius. Evidence is presented that Bledius and Eppelsheimius are sister groups. The genus has two species, E. pirazzolii and E. miricollis, that are distinguished by many characters. Both species are variable. The genus and species are described and illustrated and their distributions described. One species, E. persicus, is newly synonymized with E. pirazzolii"--P. . ItemGeneric revision of the Procirrina (Coleoptera, Staphylinidae, Paederinae, Pinophilini). (Bulletin of the American Museum of Natural History, no. 347)(American Museum of Natural History., 2010) Herman, Lee H.The paederine subtribe Procirrina is redescribed, the morphology, phylogenetic relationships, and distribution are discussed, and a key to the eight genera is presented. In the section on morphology, the female genital sclerites are discussed and terms are proposed for the fused sclerites. Neoprocirrus, Oedichirus, Oedodactylus, Palaminus, Paraprocirrus, Procirrus, and Pseudoprocirrus are redescribed and a list of the included species and examined species is included for each. Stylokyrtus, new genus, is described for Oedodactylus errans Sharp, 1876, new combination. Parapalaminus Bierig, 1943, Procirrinus Koch, 1934, and Oedichiranus Reitter, 1906, described as subgenera, are new synonyms of Palaminus, Procirrus, and Oedichirus respectively. The sister group of Procirrina is Pinophilina. The basal genus of the Procirrina is Oedodactylus; Pseudoprocirrus is the sister group to the remainder of the genera, which are clustered in two clades. Palaminus + Oedichirus comprise one of the two clades with Paraprocirrus and Procirrus + Neoprocirrus as its sister. As Stylokyrtus is known by only one specimen, most of the characters important to phylogenetic placement of the genus were unavailable for study and its position within the subtribe is ambiguous. ItemGeneric revisions of the Scopaeina and the Sphaeronina (Coleoptera: Staphylinidae: Paederinae: Lathrobiini) (Bulletin of the American Museum of Natural History, no. 460)(American Museum of Natural History., 2023-06-21) Herman, Lee H.The generic classifications of the paederine subtribes Scopaeina Mulsant and Rey, 1878, and Sphaeronina Casey, 1905, are revised. Sphaeronina, revised status, is resurrected from synonymy. Keys to the included genera of both subtribes are included. Newly discovered characters in both subtribes are discussed and illustrated. The Scopaeina now includes Scopaeus, Hyperscopaeus, Micranops, Orus, and Trisunius. The account for each genus includes its diagnostic characters, a description, summary of the general distribution, and list of the included species and specimens examined. Scopaeus Erichson, 1839, has a revised definition and is now restricted to species that have not only a constricted neck and a trichobothrium adjacent to and at about the middorsal margin of the eye, but also a metathoracic/mesofemoral stridulum comprised of a lateral, metaventral file and mesofemoral plectral ridges, slender, apically acute, metakatepisternal processes, and a middorsally fused median lobe of the aedeagus. The stridulum, redefined herein as a file and plectrum that when rubbed together produce stridulation in insects. The metaventral file and mesofemoral plectral ridges of Scopaeus, is, heretofore, unknown in the Staphylinidae or perhaps, even the Coleoptera. Variations of the stridulum and metakatepisternal processes are illustrated and described for each species group. Five genus-group names in the Western Hemisphere, Scopaeomerus Sharp, 1886, and Euscopaeus Sharp, 1886, are new synonyms of Scopaeus; Scopaeodera Casey, 1886, Scopaeoma Casey, 1905, and Scopaeopsis Casey, 1905, are revised status junior synonyms of Scopaeus. The species in those generic groups are now included in species groups of Scopaeus. Hyperscopaeus Coiffait, 1984, new status, is elevated to genus from subgeneric status in Scopaeus. Trisunius Assing, 2011, new subtribal assignment, is moved from the Medonina to the Scopaeina. Typhloscopaeus Jarrige, 1951, incertae sedis, formerly a subgenus of Scopaeus, is of unknown placement, but the species and generic names are retained in Scopaeus awaiting study of the type. Orus cervicula Casey, 1905, revised combination, is returned to Orus from Scopaeus. Orus femoralis (Sharp, 1887), new combination, is transferred from Scopaeus. There are now three named species of Orus with narrow necks. Scopaeus chiriquensis (Sharp, 1886), S. guatemalensis (Sharp, 1886), S. obscurus (Sharp, 1886), and S. palmatus (Sharp, 1886), new combinations, are transferred to Scopaeus from Scopaeomerus. Medon mexicanus (Bernhauer, 1910), new combination, is transferred to Medon from Scopaeomerus. Scopaeus crassitarsis (Sharp, 1886), S. gracilicornis (Sharp, 1886), S. impar (Bierig, 1935), new combinations, are transferred to Scopaeus from Euscopaeus.The following names are transferred from Scopaeus to Hyperscopaeus as new combinations: Hyperscopaeus admixtus (Fagel, 1973), H. albertvillensis (Fagel, 1973), H. allardianus (Fagel, 1973), H. andrewesi (Cameron, 1931), H. angolanus (Fagel, 1973), H. bamaniaensis (Fagel, 1973), H. borneensis (Cameron, 1941), H. bredoanus (Fagel, 1973), H. calidus (Bernhauer, 1932), H. confusoides (Fagel, 1973), H. confusus (Fagel, 1973), H. consimilis (Fagel, 1973), H. convexiceps (Bernhauer, 1932), H. corpulentus (Fagel, 1973), H. decelleanus (Fagel, 1973), H. dolosus (Fagel, 1973), H. endrodyanus (Fagel, 1973), H. errans (Fagel, 1973), H. erraticus (Fagel, 1973), H. fageli (Levasseur, 1981), H. fallaciosus (Fagel, 1973), H. filicornis (Fagel, 1973), H. flavidulus (Fagel, 1973), H. flavocastaneus (Lea, 1923), H. fluviatilis (Fagel, 1973), H. fossiceps (Eppelsheim, 1885), H. fuliginosus (Fagel, 1973), H. fulvescens (Motschulsky, 1858), H. fusculus (Motschulsky, 1858), H. gigantulus (Bernhauer, 1929), H. girardianus (Fagel, 1973), H. hova (Fauvel, 1905), H. hulstaertianus (Fagel, 1973), H. intermixtus (Fagel, 1973), H. kaszabianus (Fagel, 1973), H. katanganus (Fagel, 1973), H. kivuanus (Fagel, 1973), H. lamtoensis (Fagel, 1973), H. leleupianus (Fagel, 1973), H. leopoldvillensis (Fagel, 1973), H. lescuyeri (Delaunay, Coache, and Rainon, 2019), H. levasseuri (Lundgren, 1982), H. longiusculus (Fagel, 1973), H. machadoanus (Fagel, 1973), H. major (Eppelsheim, 1885), H. methneri (Bernhauer, 1932), H. minutulus (Fagel, 1973), H. mulongoensis (Fagel, 1973), H. nitidiceps (Fagel, 1973), H. nitidicollis (Fagel, 1973), H. opacicollis (Bernhauer, 1942), H. overlaetianus (Fagel, 1973), H. parvicornis (Fauvel, 1900), H. procerus (Kraatz, 1859), H. pruinosulus (Eppelsheim, 1885), H. pseudomethneri (Fagel, 1973), H. puberulus (Kraatz, 1859), H. reduncus (Fagel, 1973), H. ripicola (Fagel, 1973), H. rubricollis (Fagel, 1973), H. rubrotestaceus (Kraatz, 1859), H. ruguliceps (Fagel, 1973), H. ruziziensis (Fagel, 1973), H. semifuscus (Kraatz, 1859), H. senegalensis (Fagel, 1973), H. seydeli (Cameron, 1952), H. simillimus (Fagel, 1973), H. simulator (Fagel, 1973), H. spathiferus (Coiffait, 1970), H. spinosophallatus (Frisch, 2012), H. subconfusus (Fagel, 1973), H. subprocerus (Coiffait, 1978), H. surdus (Fagel, 1973), H. suspectus (Fauvel, 1907), H. tchapembanus (Fagel, 1973), H. thoracicus (Motschulsky, 1858), H. tristis (Bernhauer, 1929), H. vagans (Fagel, 1973, and H. voltae (Fagel, 1973). Sphaeronina, revised status, is resurrected from synonymy and now includes Sphaeronum Sharp, 1876, Tripectenopus Lea, 1918, Typhloleleupius Fagel, 1964, and Coecoscopaeus Coiffait, 1982; the last three genera are new assignments to the subtribe. Sphaeronina is redefined by the presence of a hypopharyngeal peg, an enlarged protibial concavity with combs, a ventral denticle on the left mandible, and a groove on the outer edge of the mandibles; additional possible diagnostic characters are discussed. Sphaeronum, Tripectenopus, Typhloleleupius, and Coecoscopaeus are redescribed; the genera are found, respectively, in the American tropical and subtropical regions, Australia, southern Africa and perhaps Madagascar, and Tunisia. Few African and Australian were available for study. Scopaeodracus Scheerpeltz, 1935, is a new synonym of Tripectenopus. Tripectenopus handschini (Scheerpeltz, 1935), new combination, is transferred from Scopaeodracus; Tripectenopus australiae (Fauvel, 1878), T. microps (Lea, 1923), T. pectinatrix (Lea, 1923), and T. torrensensis (Blackburn, 1891), new combinations, are transferred from Domene. ItemA new genus and new species of the Oxytelinae from Brazil (Coleoptera, Staphylinidae). American Museum novitates ; no. 2336(New York, N.Y. : American Museum of Natural History, 1968) Herman, Lee H. ItemNomenclatural changes in the Paederinae (Coleoptera, Staphylinidae). American Museum novitates ; no. 3416(New York, NY : American Museum of Natural History, 2003) Herman, Lee H.More than 70 nomenclatural changes for the Paederinae are implemented herein, and 10 problematical names that were submitted to the International Commission on Zoological Nomenclature for action are described. The spelling of the species-group names for Procirrus lefebvrei Latreille, 1829, and Achenium jejunum Erichson, 1840, are deemed the correct original spellings. The following names are new combinations. Lithocharis stilicina (Cameron, 1931) is transferred from Medon. Lobrathium alticola (Cameron, 1943), Lobrathium badium (Cameron, 1924), and Lobrathium cylindricolle (Cameron, 1924) are transferred from Lathrobium. Micranops surinamensis (Herman, 1965) and Micranops volans (Blackwelder, 1943) are transferred from Orus, and Micranops yemenicus (Coiffait, 1981) is transferred from Scopaeus. Pinobius longipennis (Cameron, 1931) is transferred from Dolicaon. Platydomene anguina (Sharp, 1874), Platydomene carinicollis (Sharp, 1889), and Platydomene funebris (Sharp, 1889) are transferred from Lathrobium, and Platydomene nobilis (Sawada, 1965) is transferred from Lobrathium. Sunius fungi (Cameron, 1943), Sunius immsi (Bernhauer, 1914), Sunius laevior (Cameron, 1943), and Sunius monticola (Cameron, 1931) are transferred from Medon. Tetartopeus cognatus (Sharp, 1889) and Tetartopeus fulvipes (Adachi, 1955) are transferred from Lathrobium. Chloecharis Lynch, 1884, is a junior synonym of Hypomedon Mulsant and Rey, 1878. The correct names for the subspecies formerly referred to as Echiaster japonicus japonicus Bernhauer, 1923 and E. j. unicolor Bernhauer, 1922, are Echiaster unicolor unicolor Bernhauer, 1922, and Echiaster unicolor japonicus Bernhauer, 1923. Lathrobium elongatulum (MacLeay, 1873) was improperly replaced and therefore is resurrected. Medon abantensis Bordoni, 1980b, is a new junior synonymic homonym of Medon abantensis Bordoni, 1980a. Medon mersinus Bordoni, 1980b, is a junior homonym of Medon mersinus Bordoni, 1980a, and a new synonym of Medon haafi Scheerpeltz, 1956. Medon siculus Coiffait, 1970, is resurrected from synonymy with Medon sicilianus Coiffait, 1970. Ochthephilum biforme (Fauvel, 1895) is resurrected from synonymy with Ochthephilum semiopacum (Eppelsheim, 1895). Paederus (Dioncopaederus) Scheerpeltz, 1957, is a new synonym of Paederus (Poederomorphus) Gautier, 1862. Platydomene abdominalis (Hubenthal, 1911) is resurrected from synonymy with Platydomene domeniformis (Koch, 1938). Panscopaeus is a valid genus, not a subgenus of Achenomorphus. Panscopaeus yakushimanus (Ito, 1992) is a new combination and is transferred from Achenomorphus. Twenty-eight junior primary homonyms are replaced. The following new names are proposed: Astenus montanus for Astenus montanellus Cameron, 1941; Astenus nepalicus for Astenus nepalensis Scheerpeltz, 1976; Homaeotarsus torquatus for Homaeotarsus collaris (Sharp, 1885); Lathrobium austellum for Lathrobium abdominale Lea, 1923; Lathrobium sinense for Lathrobium chinense Bernhauer, 1938; Lathrobium morsum for Lathrobium nanum Sharp, 1876; Lathrobium pulchrum for Lathrobium pulchellum Lea, 1923; Lithocharis chyuluensis for Lithocharis simplex Cameron, 1942; Lithocharis sordidula for Lithocharis sordida Cameron, 1928; Lobrathium rubriventris for Lobrathium rufiventre (Coiffait, 1953); Medon hatchi for Medon capitalis Hatch, 1957; Medon insulanus for Medon insularis Cameron, 1941; Medon genialis for Medon lugubris Cameron, 1932; Medon maculosus for Medon maculipennis Fagel, 1965; Medon umbrosus for Medon opacinus Scheerpeltz, 1933; Medon papuensis for Medon papuanus Cameron, 1937; Medon unicus for Medon uniformis Cameron, 1932; Mimopinophilus petilus for Mimopinophilus gracilis (Fagel, 1963); Pseudolathra lineata for Pseudolathra seriata (Sharp, 1889); Ochthephilum impavidum for Ochthephilum arrowi (Bernhauer, 1939); Ochthephilum silvestre for Ochthephilum silvaticum (Jarrige, 1970); Oreopaederus macicornis for Oreopaederus filicornis (Fagel, 1954); Palaminus lumiventris for Palaminus spiniventris Bernhauer, 1943; Palaminus trapezimorphus for Palaminus trapezicollis Bernhauer, 1918; Pinophilus infragilis for Pinophilus robustus Bernhauer, 1915; Scopaeus okei for Scopaeus gracilis Oke, 1933; Rugilus scaber for Rugilus rudis (Fagel, 1961); Scymbalium scintillans for Scymbalium nitidum Bernhauer, 1943; and Sunius impennatus for Sunius apterus (Coiffait, 1981). Tetartopeus captiosus Casey, 1905, is resurrected to replace Tetartopeus punctulatus (LeConte, 1863). Lathrobium fulvipenne (Gravenhorst, 1806) is not a junior primary homonym and should not be replaced by the next available synonym. Scopaeus okei is proposed to replace Scopaeus gracilis Oke, 1933, a junior secondary homonym. For each of the following pairs of names, the first is a nomen protectum, the second a nomen oblitum: Achenium depressum (Gravenhorst, 1802) and Achenium variegatum (Geoffroy, 1785); Astenus lyonessius (Joy, 1980) and Astenus brunneus Stephens, 1833; Domene scabricollis (Erichson, 1840) and Domene scabricollis (Heer, 1839); Lobrathium multipunctum (Gravenhorst, 1802) and Lobrathium testaceum (Paykull, 1789); Platydomene bicolor (Erichson, 1840) and Lathrobium bicolor Heer, 1839; Rugilus angustatus (Geoffroy, 1785) and Staphylinus angustatus Schrank, 1781; Rugilus geniculatus (Erichson, 1839) and Rugilus punctipennis Stephens, 1833; Rugilus rufipes Germar, 1836 and Rugilus immunis Stephens, 1833; and Scopaeus minutus Erichson, 1840 and Scopaeus pumilus (Heer, 1839). Achenium schatzmayri Koch, 1937, is currently listed with an older synonym, but that relationship requires verification. The following names were submitted to the International Commission on Zoological Nomenclature for conservation: Astenus dimidiatus (Wollaston, 1864); Astenus filum (Aubé, 1850); Astenus rutilipennis Reitter, 1909; Astenus unicolor (Mulsant and Rey, 1878); Tetartopeus fulvipes (Adachi, 1955); Lathrobium geminum Kraatz, 1857; Lathrobium pallipes Sharp, 1889; Lobrathium badium (Cameron, 1924); Paederus limnophilus Erichson, 1840; Pinophilus tenuis Fagel, 1963; and Platydomene bicolor (Erichson, 1840). ItemNomenclatural changes in the Staphylinidae (Insecta, Coleoptera). Bulletin of the AMNH ; no. 264([New York] : American Museum of Natural History, 2001) Herman, Lee H.More than 775 nomenclatural changes are proposed and nomenclatural problems are discussed for the Staphylinidae. Two generic names are proposed as replacement names: Torobus for species formerly in Trigonopselaphus, and Ryvkinius for the preoccupied Mesoporus Ryvkin. Type species are fixed or problems with the type species are discussed for: Entomoculia, Eumegalopsidia (an unavailable name), Anthobium, Lesteva, Phyllodrepoidea, Aleioglyphesthus, Bolitogyrus, Cephalonthus, Diatrechus, Euremus, Indoquedius, Philothalpus, Philonthopsis, Pseudoremus, Leptophius, Stenus (Nestus), Coproporus, and Paratachinus. The type species for Entomoculia, Anthobium, Phyllodrepoidea, Leptophius, and Stenus (Nestus) are fixed under provisions of article 70.3 of the Code. Emendations are cited or one of multiple original spellings adopted in Bledius, Bryoporus, Carpelimus, Edaphus, Eleusis, Gauropterus, Indosorius, Lispinus, Lithocharodes, Medon, Osorius, Philonthus, Piestus, Pinophilinus, Platydracus, Priochirus, Prognathoides, Quedius, Stenus, and Trigonuras. New combinations are proposed in Arrhenopeplus (3), Dialycera (2), Eusphalerum (14), Nacaeus (19), Pseudoxyporus (4), Homalotrichus (1), Thinodromus (2), Gabrius (1), Platydracus (57), Torobus (9), Bryophacis (7), Ischnosoma (16), Lordithon (66), Ryvkinius (1), and Sepedophilus (244). Fifty-nine new synonyms are listed for Micropeplus (1), Anthophagus (2), Eusphalerum (1), Phloeonomus (1), Eleusis (1), Leptochirus (1), Neolosus (1), Osorius (5), Priochirus (1), Anotylus (1), Bledius (1), Carpelimus (1), Ochthephilus (1), Apoquedius (1), Diatrechus (1), Erichsonius (2), Gabrius (2), Gastrisus (1), Hesperus (2), Heterothops (2), Leptacinus (1), Notolinus (1), Phallolinus (1), Philonthus (6), Quedius (5), Staphylinus (2), Tasgius (1) Xantholinus (4), Stenus (7), Sepedophilus (1), and Tachinus (1). Polyphematiana E. Strand is a junior synonym of Trigonopselaphus Gemminger and Harold. Eleven names are resurrected from synonymy, one each in Phyllodrepoidea, Anotylus, Oxytelus, Loncovilius, Philonthus, Platydracus, Lordithon, Mycetoporus, and Tachinus and two in Stenus. Under provisions of article 23.9.1 of the Code, 28 junior synonyms are protected in Amphichroum (1), Carpelimus (1), Deleaster (1), Proteinus (1), Bisnius (1), Leptacinus (1), Megalinus (1), Neobisnius (1), Ocypus (1), Philonthus (3), Quedius (4), Tasgius (2), Stenus (5), Bolitobius (1), Lordithon (1), Mycetoporus (1), and Tachyporus (2). Provisions of the same article protect 5 junior homonyms in Eusphalerum (1), Xylodromus (1), Quedius (1), Tachinus (1), and Tachyporus (1). Under provisions of article 23.9.3 of the Code, 15 junior synonyms will be referred to the Commission for rulings under the plenary power. Pending the outcome of these cases, use of the junior name is maintained. The affected species are in Eusphalerum (1), Lesteva (1), Omalium (1), Phloeostiba (1), Xylodromus (1), Anotylus (1), Bledius (1), Carpelimus (1), Ocypus (1), Philonthus (1), Quedius (4), and Lordithon (1). Under provisions of article 23.9.5 of the Code, 64 junior primary homonyms will be referred to the Commission for rulings under the plenary power. Pending the outcome of these deliberations, use of the junior name is maintained. The species are in Eusphalerum (1), Mannerheimia (1), Omaliomimus (1), Omaliopsis (1), Omalium (4), Phyllodrepa (1), Pycnoglypta (1), Xylodromus (1), Anotylus (1), Bledius (1), Carpelimus (1), Oxyporus (1), Belonuchus (2), Bisnius (1), Cafius (2), Cheilocolpus (1), Diatrechus (1), Endeius (1), Gabrius (5), Hesperus (2), Leptacinus (1), Nordus (1), Paederomimus (1), Philonthus (10), Platydracus (3), Quedius (2), Staphylinus (6), Xantholinus (2), Xanthopygus (1), Xenopygus (2), Carphacis (1), Coproporus (1), Tachinomorphus (1), and Tachinus (3). Replacement names are proposed for 90 preoccupied names in 37 genera: Euaesthetus (1), Leptotyphlus (1), Micropeplus (1), Eusphalerum (2), Lesteva (1), Olophrum (1), Omalium (2), Clavilispinus (1), Eleusis (3), Holotrochus (3), Leptochirus (1), Lispinus (4), Osorius (3), Priochirus (2), Thoracochirus (1), Anotylus (2), Carpelimus (2), Platystethus (1), Atanygnathus (1), Bisnius (1), Diochus (1), Gabrius (3), Hesperus (2), Heterothops (1), Leptacinus (3), Philonthus (19), Quedius (4), Staphylinus (3), Tasgius (1), Dianous (1), Bolitobius (1), Carphacis (1), Coproporus (3), Lordithon (3), Sepedophilus (5), Tachinus (4), Tachyporus (1). Eight names are resurrected to replace preoccupied names in Anotylus (1), Philonthus (1), Quedius (1), Stenus (2), Lordithon (1), Tachinomorphus (1), and Tachinus (1). Twenty-six senior homonyms, in Hapalaraea (1), Omalium (2), Phyllodrepa (1), Bledius (1), Oxytelus (1), Bisnius (1), Quedius (1), Staphylinus (15), Carphacis (1), Sepedophilus (1), and Tachinus (1), are regarded to be "nomina dubia" and the junior names conserved by fiat, without sanction by the Code. Psephidonus is the older name, but application to the Commission will be made to reject that name in favor of the younger Geodromicus. Tachyporiniformes is an unavailable name. ItemPhylogeny and reclassification of the genera of the rove-beetle subfamily Oxytelinae of the world (Coleoptera, Staphylinidae). Bulletin of the AMNH ; v. 142, article 5(New York : [American Museum of Natural History], 1970) Herman, Lee H. ItemPseudopsis : two new species from India (Coleoptera, Staphylinidae, Pseudopsinae). American Museum novitates ; no. 2755(New York, N.Y. : American Museum of Natural History, 1983) Herman, Lee H."Two new species of Pseudopsis (susae and suchi) from India are described and illustrated. They are part of the sulcata complex which is represented in the New World by 19 species and, until now, in the Old World by five, only two of them from India. It is observed that further collecting will produce more species of the complex. A key to the eight Old World species of Pseudopsis is provided"--P. . ItemRevision and phylogeny of the monogeneric subfamily Pseudopsinae for the world (Staphylinidae, Coleoptera). Bulletin of the AMNH ; v. 155, article 3(New York : American Museum of Natural History, 1975) Herman, Lee H."Pseudopsis is redescribed, a key to the species is presented, and the phylogeny, distributional history, and biology of the species are discussed. Of the 30 species in the genus five are redescribed, 24 are newly described, and one, P. columbica, is resurrected from synonymy with P. sulcata, but was not studied. Pseudopsis montoraria is proposed to replace what mistakenly has been called P. obliterata. The true P. obliterata and P. detrita are synonyms, the former taking priority. What was previously the geographically widespread P. sulcata is a complex of 24 sibling species. The new species of the P. sulcata complex in the Old World are P. afra, P. himalayensis, P. prolixa, and P. watanabei. In the new World the complex is represented by the newly described P. subulata, P. sagitta, P. obtusa, P. callosa, P. abbreviata, P. sinuata, P. echinata, P. maja, P. spicula, P. bilacuna, P. biloba, P. constricta, P. vespina, P. lata, P. dybasi, P. petila, P. dilata, P. grossa, and P. wygodzinskyi. Pseudopsis sulcata (sensu stricto) occurs in the western Palaearctic region. Most species of the complex are allopatric, but nine occur sympatrically, three at one locality, and two at each of three other places. Pseudopsis is the only genus of the Pseudopsinae and is the sister group of the Oxytelinae. Two subgenera are recognized: Pseudopsis and Pseudopsiella. Chiliopseudopsis is a junior synonym of Pseudopsiella. The nominal subgenus contains the 25 species of the sulcata complex. Pseudopsis obliterata, P. montoraria, P. minuta, P. arrowi, and P. adustipennis comprise Pseudopsiella. Pseudopsis and Pseudopsiella are sister groups. Within Pseudopsiella, P. minuta is the sister species of the other four species. Pseudopsis obliterata and P. montoraria are the sister group to P. arrowi and P. adustipennis. Alternative phylogenetic schemes are discussed. The species of the genus live principally in temperate, montane regions of the world. They probably originated in western North America and spread from there"--P. 245. ItemRevision and phylogeny of Zalobius, Asemobius, and Nanobius, new genus (Coleoptera, Staphylinidae, Piestinae). Bulletin of the AMNH ; v. 159, article 2(New York : American Museum of Natural History, 1977) Herman, Lee H."Zalobius and Asemobius are redescribed, Nanobius is newly described and their phylogeny and taxonomic position are discussed. One new species, nancyae, is descibed in Zalobius, and serricollis, formerly in Zalobius, is placed in the new genus Nanobius. The four species of the three genera are keyed, described, and illustrated. The current subfamilial assignement of the genera is concluded to be unsatisfactory but the genera share no detected derived characters with other subfamilies that permit reassignment. Cladistic analysis indicates that the two species of Zalobius form the sister group of Asemobius caelatus, that Nanobius serricollis is the sister species of a group containing all three, and that the four species share at least six apomorphic character states. Alternative phylogenetic schemes are discussed. The four species live in the temperate, montane areas of the western United States and Canada"--p. 49. ItemRevision of Bledius and related genera. Part 1, The aequatorialis, mandibularis, and semiferrugineus groups and two new genera (Coleoptera, Staphylinidae, Oxytelinae). Bulletin of the AMNH ; v. 149, article 2(New York : [American Museum of Natural History], 1972) Herman, Lee H."Bledius is redescribed, and a key to the species-groups of the New World is presented. The species of the aequatorialis, mandibularis, and semiferrugineus groups of Bledius Leach from the Nearctic and West Indian areas and two new genera, Microbledius and Psamathobledius, are described. Illustrations, keys, diagnoses, and distributional and habitat data are presented for each species. Three new species, Microbledius playanus, Microbledius actitus, and Microbledius litoreus are described and M. forcipatus (Le Conte), M. albidus (Sharp), M. albipennis (Bernhauer), M. bifasciatus (Sharp), M. miles (Bernhauer), M. minutissimus (Bernhauer), and M. weiseri (Bernhauer) are transferred from Bledius. Psamathobledius contains P. caribbeanus (Blackwelder), P. microcephalus (Fauvel) and P. punctatissimus (Le Conte) (B. esposus Blackwelder is a new synonymy); all are transferred from Bledius. Bledius aequatorialis Mutchler, B. beattyi Blackwelder, and B. ceratus Blackwelder are included in the aequatorialis group. Bledius mandibularis Erichson, B. pallipennis Say (B. gularis Le Conte is a new synonymy), B. fortis Le Conte, B. ferratus Le Conte, and B. jacobinus Le Conte (B. actus Herman is a new synonymy) are in the mandibularis group. Bledius semiferrugineus Le Conte (B. canaliculatus Notman is a new synonymy), B. foraminosus Casey (B. relictus Fall and B. deceptivus Fall are new synonymies), B. tallaci Fall, B. gravidus Casey (B. missionensis Hatch is a new synonymy), B. rotundicollis Le Conte, B. rubiginosus Erichson, B. fumatus Le Conte, B. philadelphicus Fall (B. dickersoni Notman and B. mixtus Notman are new synonymies), and B. analis Le Conte, B. nitidicollis Le Conte, B. assimilis Casey, and B. coulteri Hatch are included in the semiferrugineus group"--P. 115. ItemRevision of Bledius and related genera. Part 2, The armatus, basalis, and melanocephalus groups (Coleoptera, Staphylinidae, Oxytelinae). Bulletin of the AMNH ; v. 157, article 2(New York : American Museum of Natural History, 1976) Herman, Lee H."In the present paper, the second of a three part series on Bledius, the armatus, basalis, and melanocephalus groups from the Nearctic region are discussed. In addition to a revised key to the species groups of the New World, illustrations, keys, diagnoses and distributional and habitat data are presented for each species. Bledius notialus, B. aquilonarius, and B. thinopus are newly described. Bledius cuspidatus Le Conte, B. agonus Casey, B. tenuis Casey, and B. nelsoni Hatch are new junior synonyms of B. flavipennis Le Conte. Bledius lectus Casey, B. piceus Fall, B. specularis Fall, B. regularis Fall, and B. condonensis Hatch are new junior synonyms of B. opacifrons Le Conte. Bledius nigriceps Notman is a new junior synonym of B. politus Erichson. Bledius gradatus Fall and B. boddyi Hatch are new junior synonyms of B. eximius Casey. Bledius arizonensis Fall is a new junior synonym of B. strenuus Casey. Bledius ignavus Casey and B. misellus Casey are new junior synonyms of B. basalis Le Conte. Bledius confinus Fall is a new junior synonym of B. melanocephalus (Say). Bledius melanocephalus (Say), the only species in the melanocephalus group, is recognized and discussed for the first time in more than 150 years. Bledius cordatus (Say), B. basalis Le Conte, B. turbulentus Casey, B. dimidiatus Le Conte, B. opaculus Le Conte, B. neglectus Casey, and B. thinopus, new species, comprise the basalis group and, but for B. dimidiatus, all live on the coast of eastern North America. The armatus group includes B. flavipennis Le Conte, B. nitidiceps Le Conte, B. opacifrons Le Conte, B. ineptus Casey, B. consimilis Fall, B. monstratus Casey, B. fenyesi Bernhauer and Schubert, B. politus Erichson, B. episcopalis Fall, B. eximius Casey, B. bellicus Blackwelder, B. strenuus Casey, B. aquilonarius, new species, and B. notialus, new species. All species of the armatus group except B. politus live principally west of the Mississippi River"--P. 75. ItemRevision of Bledius. Part 3, The annularis and emarginatus groups (Coleoptera, Staphylinidae, Oxytelinae). Bulletin of the AMNH ; v. 175, article 1([New York] : American Museum of Natural History, 1983) Herman, Lee H."In the present report, the third in a series on Bledius, the annularis and emarginatus groups are discussed and a new species is described in the aequatorialis group. A key to the species, descriptions, diagnoses, illustrations, and distributional and habitat data for each species are presented. Bledius habrus, melanocolus, aurantius, jucundus, naius, nardus, omega, venus, viriosus, wudus, and zophus are newly described. Thirteen nominal species are recognized as synonyms. Bledius albidipennis Bernhauer and ornatus LeConte are new junior synonyms of albonotatus Mäklin; apicalis Fall is a new junior synonym of diagonalis LeConte; adjustus Casey and transitus Fall are new junior synonyms of gentilis Casey; kincaidi Hatch is a new junior synonym of parvicollis Casey; rusticus Fall is a new junior synonym of bicolor Casey; luteipennis LeConte, medialis Fall, oregonensis Hatch, and pleuralis LeConte are new junior synonyms of suturalis LeConte; and borealis Blatchley and bowronensis Hatch are new junior synonyms of turgidus Casey. The species in the United States included in the emarginatus group are cognatus LeConte, emarginatus (Say), and wudus Herman. They are found in the eastern and southeastern United States. The species are similar and not easily distinguished. Bledius albonotatus Mäklin, aurantius Herman, bicolor Casey, cedarensis Hatch, confusus LeConte, diagonalis LeConte, gentilis Casey, gracilis Casey, habrus Herman, jucundus Herman, laticollis LeConte, melanocolus Herman, monticola Casey, naius Herman, nardus Herman, newelli Hatch, omega Herman, parvicollis Casey, persimilis Fall, phytosinus LeConte, ruficornis LeConte, suturalis LeConte, tarandus Herman, tau LeConte, turgidus Casey, venus Herman, villosus Casey, viriosus Herman and zophus Herman are species from the United States and Canada included in the annularis group and which I can identify. Also included in the annularis group are nine species listed together in what I call the annularis complex: annularis LeConte, breretoni Hatch, honestus Casey, languidus Casey, mysticus Fall, nebulosus Casey, sinuatus LeConte, stabilis Casey, and washingtonensis Hatch. After years of study and collecting I have been unable to resolve the species of this complex which is widespread in Canada and the northern United States. I was unable to resolve three other species: fasciatus (Say), longipennis Mäklin, and verticalis Notman; the types of each have been lost but I discuss each of them. The annularis group species are widespread in the United States and Canada and most are found in shaded soil near fresh water. Only two, albonotatus and newelli, are associated with coastal, saline habitats. Several species, notably ruficornis, tarandus, and turgidus, have transcontinental distributions, and one, albonotatus, is found along the coast from western Alaska to Baja California. The addition of a new species, susae, to the aequatorialis group brings that group up to four. However, susae is the only one of the group known only from the United States which is found only on the mainland. It is also the sole species of Bledius in which the females have a pronotal horn; although pronotal horns are known in a number of New and Old World species, heretofore no females have been reported with such a modification. Since publication of Parts I and II of this monograph, I have examined over 6000 more specimens of 41 previously revised species. In Appendix II all the new records are included; for many species extensions of their previously known range are recorded. Lectotypes are designated for the species of Bledius described by LeConte, Casey, and Fall. For Part III, 28,408 specimens were examined"--P. 4. ItemRevision of Bledius. Part 4, Classification of species groups, phylogeny, natural history, and catalogue (Coleoptera, Staphylinidae, Oxytelinae). Bulletin of the AMNH ; v. 184, article 1([New York] : American Museum of Natural History, 1986) Herman, Lee H."The purpose of this monograph is to review and reinterpret what is known about Bledius with the goal of prompting further study of the genus. Included are a new infrageneric classification for the world; discussions of the life cycle, habitat, distribution, enemies, chemical secretions, economic importance, immature stages, fossils, and phylogeny; and a cross-indexed annotated catalogue to the taxonomic names and associated literature. Formal taxonomic changes include the following: Bledius minniensis from Armenia is a new species, Neobledius is a new synonym of Bledius and Elbidus, and Microbledius and Psamathobledius are new synonyms of Bledius. New combinations include B. karachiensis transferred from Neobledius and actitus, litoreus, and playanus transferred from Microbledius. New names include albanicus and jutlandensis which replace the preoccupied nebulosus Koch and atlanticus Lohse, respectively. The following names are labeled as new status: B. castaneipennis Mannerheim is elevated from synonymy with opacus Block; atramentarius Rottenberg is elevated from synonymy with and replaces bos Fauvel which was described a year later; limicola Tottenham is elevated from synonymy with and replaces germanicus Wagner which is preoccupied by germanicus Gravenhorst; minor Mulsant and Rey is elevated from synonymy with and replaces devillei Bondroit which was an unnecessary replacement name. Two names are emended: cariniceps for carinlceps and ghesquierei for guesquierei. Six names, bubalus Gistel, castaneus Mulsant and Rey, chimerinus Gistel, germanicus Gravenhorst, gyllenhalii Laporte, and westerhauseri Gistel are used herein for the first time since their original publication. All eight currently recognized subgenera are treated as synonyms of Bledius. In their place a classification of 34 species groups is proposed. This classification is based on examination of 422 of the 439 species of Bledius. The species were assigned to species groups based on examination of type material or subsequently identified specimens. Nine species were assigned to species groups from characters in the original description -- each such assignment is clearly indicated. Eight species are listed as incertae sedis for one or more of the following reasons: material was unavailable for study, the description did not cite sufficient information for placement in a species group, the types were lost, or the species was unknown. Descriptions, illustrations, a list of included species and localities, and a distributional map are provided for each species group along with a key to the group. No effort was made to define or revise species. The monophyly of Bledius is established by the presence of a cluster of secretory pores in a prosternal pit or depression and the possession in females of a pair of elongate, undivided genital sclerites on segment IX. Both Bledius and its sister genus Eppelsheimius have a central row of spine-like setae on the hypopharynx; they share other characters as well. The relationships among the species groups are shown in a cladogram on which many characters are homoplasic. Bledius comprises two main lineages. One with 14 species groups and 88 species includes, among others, the large robust species with horns. Most species of this branch live in saline habitats. Two of the species groups are restricted to the New World. The sister lineage includes the remaining 20 species groups and more than 340 species. A terminal cluster of species groups along with species in three other groups live in saline habitats; the others, including the four most speciose groups, live in freshwater habitats. In this lineage 3 species groups are confined to the New World, 11 to the Old, and the remaining 6 are widespread in both hemispheres. The primitive habitat for Bledius was near salt water. One descendent remains there; the other adapted to the freshwater habitat. In this second lineage some highly speciose groups developed near freshwater and there was a reinvasion of the saline habitat. Bledius is a large genus of 439 species, some of which occur in huge populations. Species are found on all continents except Antarctica and on most continental islands but are absent from most oceanic islands. Although only a third of the species lives in saline habitats (inland as well as coastal), these species represent two-thirds of the species groups. Two-thirds of the species live in freshwater habitats, representing the remaining third of the species groups. Bledius may have lived as far back as the Cretaceous. Nine fossil species have been described as belonging in Bledius, however, two of these are no longer included. Of the remaining seven only Bledius glaciatus from the Pleistocene and B. primitiarum of the Oligocene are likely to be Bledius. Another probable member of Bledius is Staphylinus lesleyi Scudder also of the Oligocene; this species has not been moved to Bledius. Although the oldest probable fossils of Bledius are about 35 million years old, evidence suggests that the Oxytelinae existed about 150 million years ago during the Jurassic. Species of Bledius are most abundant in unvegetated or lightly vegetated, sunny, moist sand adjacent to rivers, lakes, and oceans. They preferentially select their habitat based on soil moisture, salinity, texture, and, indirectly at least, size of the sand grains, availability of food, and amount of shade. The pressure of predators may influence habitat limits but that may be partly due to the physiological weakening of a Bledius that is not living in its optimal habitat. Although as yet undemonstrated, food plant specificity may influence habitat selection by Bledius. Bledius lives in burrows which adults excavate with their mandibles and strengthen with their protibiae. The burrows are several millimeters to about 40 cm deep and about 1 to 5 mm in diameter. Larvae also construct and live in burrows. Eggs are laid in special egg chambers; some species cache food in the burrow. The burrow serves not only as living quarters for Bledius but helps protect them. Bledius can live at a relative humidity of 100 percent to no less than 93 percent. The humidity within the burrow is nearly 100 percent. Some seacoastal species live in the intertidal zone where they remain during high tide. Individuals that stay in the burrow during high tide live in an air bubble, which acts as a physical gill, and are immediately active after the tide recedes. Individuals exposed directly to seawater fly away or survive by becoming comatose and recover only slowly when removed from the water; most, however, die after a few hours of immersion. Usually the adults overwinter in deep burrows. Eggs are deposited during spring or early summer and develop in two to three weeks. The larval stage lasts for six to eight weeks and the five instars are each of undetermined length. Pupae exist for about a week. Egg to adult then is two to three months. During the summer adults live for three to six weeks. Females oviposit several times; the number of generations per year is unclear but may vary according to species and climate. Larval characters have been published for 19 species. Adults and larvae feed on algae and diatoms that live in the moisture surrounding each sand grain. The green algae there are said to be stunted filamentous forms. Some Bledius facultatively store algae. Some species are said to exhibit subsocial behavior by attending to their offspring. Evidence for this interpretation is inconclusive. Carabid beetles of Dyschirius are regarded as the principal predators of Bledius. However, little evidence is found to support a species-specific host-prey relationship. More likely, the relationship of Bledius and Dyschirius is based on geographical distribution and habitat preference. Over 50 species of Dyschirius are reported with over 80 species of Bledius. Other Carabidae reported as actual or probable predators of Bledius include species of Cillenum, Bembidion, Pogonus, Dicheirotrichus, Clivina, and Schizogenius. The ichneumonid wasp, Barycnemis blediator, is a parasitoid of Bledius spectabilis. Sandpipers and rainbow trout also eat Bledius. Fungi of the order Laboulbeniales commonly infest Bledius and, although usually regarded as benign, one report suggests otherwise. Species of Bledius secrete a characteristic, penetratingly fragrant chemical of five components. That this secretion functions as a defensive chemical is debatable. No experimentally supported hypotheses have explained the function of this compound. One species in Japan was reported to do economically significant damage to clay beds used to produce salt by evaporation of seawater. The literature for and taxonomic names used in Bledius are summarized in a catalogue. Fifteen generic-level names have been used in Bledius; 646 species-level names have been used, 434 of which are presently listed as valid and extant. All names that have been included in Bledius are listed in the catalogue. The species names are cross-indexed and the references annotated"--P. 5-6. ItemRevision of Stereocephalus (Coleoptera, Staphylinidae, Paederinae). American Museum novitates ; no. 2683(New York, N.Y. : The American Museum of Natural History, 1979) Herman, Lee H."Stereocephalus, which occurs east of the Andes from Venezuela to northern Argentina, is transferred from one paederine subtribe, the Dolicaonina, to another, the Lathrobiina. Three new species, S. myrigeus, S. ruhus, and S. rinnanus, all from Brazil, are described. Stereocephalus seriatipennis Lynch is redescribed and S. dilaticeps Bernhauer is placed as a junior synonym of seriatipennis. A key and illustrations are provided. Stereocephalus seriatipennis and S. rinnanus are sister species and together with S. ruhus comprise the sister group of S. myrigeus"--P. . ItemRevision of the Asian tribe Megarthropsini (Coleoptera, Staphylinidae, Tachyporinae). American Museum novitates ; no. 3430(New York, NY : American Museum of Natural History, 2004) Herman, Lee H.The species and genera of the Megarthropsini are revised. Descriptions and illustrations of the genera and species and keys for identification are presented. Lacvietina, a new genus, with four new species is described. Three of the new species, L. aurora, L. copiosa, and L. cuprina, are from Vietnam, and the fourth, L. paricosta, is from Malaysia and Thailand. Lacvietina punctatissima (Hayashi), from Taiwan, is a new combination and was transferred from Tachinus. A new species of Nepaliodes, N. solangelae from Thailand and China, is described. Seven new species of Megarthropsis are described; six of them, M. deverra, M. durga, M. empusa, M. frazerensis, M. parca, and M. smetanai, are from Malaysia; the seventh, M. djawaensis, is from Indonesia. A phylogenetic analysis supports Megarthropsini as a monophyletic tribe with Deropini as its sister group. Within Megarthropsini, Peitawopsis is the most basal genus to the remaining three genera, and Lacvietina is the sister group to Megarthropsis and Nepaliodes. The tribe is most readily distinguished from other taxa of the Tachyporinae by the densely and coarsely punctate body. Finally, the type species of Paratachinus, P. laticollis Cameron, is redescribed and discussed. Lacvietina takashii is a new combination. ItemRevision of the New World species of Oedichirus (Coleoptera, Staphylinidae, Paederinae, Pinophilini, Procirrina). (Bulletin of the American Museum of Natural History, no. 375)(American Museum of Natural History., 2013-03-15) Herman, Lee H.The New World species of Oedichirus Erichson, 1839, are revised, redescribed, or newly described, illustrated, and included in keys for identification. The morphology of the ventral pterothoracic sclerites is examined and Matsuda's (1970) interpretation is largely adopted over that of Ferris (1940b). Newly discovered characters of the median gonocoxal plate and associated vulvar plate, including its microstructures, are discussed. Prior to the present paper New World species were known in Brazil and Costa Rica, but are herein newly reported for the Dominican Republic, Mexico, Ecuador, Peru, Bolivia, and Argentina; a specimen intercepted at a port in the United States was said to have come from Nicaragua. Eight species were previously described; 19 new ones are added herein, 12 from Brazil (O. apiculus, O. batillus, O. bicristatus, O. bullaglaber, O. bullahirtus, O. clavolateralis, O. clavulus, O. echinatus, O. exilis, O. glabrihamus, O. lunatus, and O. procerus), two from Mexico (O. isthmus and O. sinuosus), and one each from Argentina (O. misionesiensis), Bolivia (O. dilophus), Ecuador (O. distortus), Peru (O. hamatus), and Dominican Republic (O. dominicanensis). A neotype is designated for O. geniculatus and lectotypes are designated for O. brunneus Wendeler, O. pictipes Bierig, O. ohausi Wendeler, O. optatus Sharp, O. sparsipennis Bernhauer, and O. speculifrons Bernhauer. Some species are included in species groups. ItemA revision of the rove-beetle genus Charhyphus (Coleoptera, Staphylinidae, Phloeocharinae). American Museum novitates ; no. 2496(New York, N.Y. : American Museum of Natural History, 1972) Herman, Lee H."Trigites Handlirsch, 1907, and Siberia Blackwelder, 1952, are placed as junior synonyms of Charhyphus Sharp, 1887, and the genus and its species redescribed, illustrated, placed into a key and their phenetic relationships and biology discussed. Charhyphus arizonensis is newly described. The position of the fossil species, Charhyphus coeni, is not certain. Charhyphus should be in the Phloeocharinae, not the Piestinae where some of the species had previously been placed"--P. .