Browsing by Author "Saucedo, Alma D."
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Item The goblin spiders of the new genus Volborattella (Araneae, Oonopidae) from Madagascar. (American Museum novitates, no. 3822)(American Museum of Natural History., 2015-01-16) Saucedo, Alma D.; Ubick, Darrell.; Griswold, Charles E.; Goblin Spider Planetary Biodiversity Inventory.A new genus of goblin spider from Madagascar, Volborattella Saucedo and Ubick, is proposed and its five included species newly described and illustrated: V. teresae, the type species, V. guenevera, V. nasario, V. paulyi, and V. toliara. These species differ from other oonopids in several unusual characters, especially the variously modified setae: abdominal scutes having thick recumbent setae with large bases and conspicuous pits; the pedicel region with mats of plumose setae and associated cuticular projections; and anterior metatarsi with prolateral combs. The male palp of Volborattella appears to be unique in having a terminal projection (embolar superior prong, ESP) that forms an abrupt spiral and the female a receptaculum with an accessory duct (curved tube). Volborattella resemble members of the Gamasomorpha complex in lacking leg spines and having a flattened abdomen with complete scutes, but differ genitalically. The Volborattella female has a receptaculum that is wider than long (as opposed to longer than wide in the Gamasomorpha complex) and the male has the embolar region sharply bent (as opposed to evenly curved), which places the genus in the Pelicinus complex. The relationship of Volborattella to other pelicinoids is not resolved. Although the genus most closely resembles some Silhouettella Benoit, Noideattella Álvarez-Padilla et al. and Lionneta Benoit in various genitalic features, somatically it shares with Tolegnaro Álvarez-Padilla et al. and two undescribed Malagasy genera the gamasomorphoid body form and plumose setae. Volborattella females are unusual in having antisymmetrical internal genitalia, with the CTs occurring in either a left or right position, and that these differences in handedness (antisymmetry) are evenly distributed. The shape and size of the CT, as examined through light and scanning microscopy, suggest that it is a coupling pocket for the ESP; if so, Volborattella females are either righties or lefties in their palpal preference.Item Tarsal organ morphology and the phylogeny of goblin spiders (Araneae, Oonopidae), with notes on basal genera. (American Museum novitates, no. 3736)(American Museum of Natural History., 2012-02-29) Platnick, Norman I.; Abrahim, Naiara.; Alvarez-Padilla, Fernando.; Andriamalala, Daniela.; Baehr, Barbara, 1953-; Baert, L.; Bonaldo, Alexandre B.; Brescovit, Antonio D.; Chousou-Polydouri, Natalia.; Dupérré, N. (Nadine); Eichenberger, Beata.; Fannes, Wouter.; Gaublomme, Eva.; Gillespie, Rosemary G., 1957-; Grismado, Cristian J.; Griswold, Charles E.; Harvey, Mark S.; Henrard, Arnaud.; Hormiga, Gustavo.; Izquierdo, Matías A.; Jocqué, R.; Kranz-Baltensperger, Yvonne.; Kropf, Christian.; Ott, Ricardo.; Ramírez, Martín J.; Raven, Robert J.; Rheims, Cristina A.; Ruiz, Gustavo R. S.; Santos, Adalberto J.; Saucedo, Alma D.; Sierwald, Petra.; Szűts, Tamás.; Ubick, Darrell.; Wang, Xin-Ping.Based on a survey of a wide variety of oonopid genera and outgroups, we hypothesize new synapomorphies uniting the Oonopidae (minus the South African genus Calculus Purcell, which is transferred to the Orsolobidae). The groundplan of the tarsal organ in Oonopidae is hypothesized to be an exposed organ with a distinctive, longitudinal ridge originating from the proximal end of the organ, and a serially dimorphic pattern of 4-4-3-3 raised receptors on legs I-IV, respectively. Such organs typify the diverse, basal, and ancient genus Orchestina Simon. Several other genera whose members resemble Orchestina in retaining two plesiomorphic features (an H-shaped, transverse eye arrangement and a heavily sclerotized, thick-walled sperm duct within the male palp) are united by having tarsal organs that are partly (in the case of Cortestina Knoflach) or fully capsulate (in the case of Sulsula Simon, Xiombarg Brignoli, and Unicorn Platnick and Brescovit). The remaining oonopids are united by the loss of the heavily sclerotized palpal sperm duct, presumably reflecting a significant transformation in palpal mechanics. Within that large assemblage, a 4-4-3-3 tarsal organ receptor pattern and an H-shaped eye arrangement seem to be retained only in the New Zealand genus Kapitia Forster; the remaining genera are apparently united by a reduction in the tarsal organ pattern to 3-3-2-2 raised receptors on legs I-IV and by the acquisition of a clumped eye arrangement. Three subfamilies of oonopids are recognized: Orchestininae Chamberlin and Ivie (containing only Orchestina; Ferchestina Saaristo and Marusik is placed as a junior synonym of Orchestina), Sulsulinae, new subfamily (containing Sulsula, Xiombarg, Unicorn, and Cortestina), and Oonopinae Simon (containing all the remaining genera, including those previously placed in the Gamasomorphinae). The type species of Sulsula and Kapitia, S. pauper (O. P.-Cambridge) and K. obscura Forster, are redescribed, and the female of S. pauper is described for the first time. A new sulsuline genus, Dalmasula, is established for Sulsula parvimana Simon and four new species from Namibia and South Africa.